1.09.1960: Date of Birth
1980 - 1987: Studies in Biology and German Literature at the University of Heidelberg, Germany.
1990: PhD (University of Heidelberg)
1990 - 1992:Postdoctoral work at LONZA Ltd, Basle and Visp, Switzerland
1992 - 1994: Research Scientist at the University of Hohenheim, Germany
1995 - today: group leader at the University of Tübingen, Germany.
1999: Habilitation (German Associate Degree)
2000 - present: Head of the Steinbeis-Transfer Center for Ecotoxicology and Ecophysiology, Rottenburg, and head of the histology group at the Animal Physiology, University of Tübingen
since 2006: Apl. Prof. at the University of Tuebingen
biomonitoring, assessment of environmental stress, physiological ecology of fish and invertebrates